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Her first audition was for the NBC police drama Homicide: Life on the Street.
She did not get the role but knew from then on that she just wanted to act.
Her mother's side of the family has held ownership in the NFL's Pittsburgh Steelers football team since its formation in 1933, and her father's side of the family has held ownership in the NFL's New York Giants since its formation in 1925.
Her paternal grandparents were Wellington Mara and Ann Mara.
She is the second of four siblings, with one older brother, Daniel; one younger sister, Patricia (known professionally as Rooney Mara, also an actress); and one younger brother, Conor.
She made her film debut in 1999 with Random Hearts.Mara also appeared on Cold Case, Boston Public, CSI: Miami and CSI: Crime Scene Investigation that same year.Mara was cast as the lead in the 2004 The WB pilot Prodigy, about a teenage child prodigy.On July 10, 2014, she was nominated for the Primetime Emmy Award for Outstanding Guest Actress in a Drama Series.She made her stage debut in 2003 at the Williamstown Theatre Festival in John Guare's Landscape of the Body with Lili Taylor.